Anatomy of Flowering Plants
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Class: 11 Biology
Chapter: Anatomy of Flowering Plants (MP Board)
1. Introduction & The Tissues
Anatomy is the study of internal structures of plants. A tissue is a group of cells having a common origin and usually performing a common function.
A. Meristematic Tissues (विज्योतक ऊतक)
Cells have the capacity to divide actively.
- Apical Meristems: Present at root and shoot apices; responsible for primary growth (increase in length).
- Intercalary Meristems: Located between mature tissues; occurs in grasses and helps regenerate parts removed by grazing.
- Lateral Meristems (Secondary Meristems): Cylindrical meristems (e.g., Fascicular cambium, Cork cambium); responsible for secondary growth (increase in girth/thickness).
B. Permanent Tissues (स्थायी ऊतक)
Cells have lost the capacity to divide and have attained maturity.
-
Simple Permanent Tissues (सरल स्थायी ऊतक): Made of only one type of cells.
- Parenchyma: Living, thin-walled cells. Functions: Photosynthesis, storage, secretion.
- Collenchyma: Living, unevenly thickened corners (pectin deposition). Function: Mechanical support to growing parts (young stem, petiole).
- Sclerenchyma: Dead, highly thick walls (lignified). Function: Mechanical support. Divided into fibers and sclereids (stone cells).
-
Complex Permanent Tissues (जटिल स्थायी ऊतक): Made of more than one type of cells working as a unit.
- Xylem (दारू): Conducts water and minerals from roots to stem and leaves.
- Elements: Tracheids, Vessels, Xylem parenchyma (living), Xylem fibers (dead).
- Note: Gymnosperms lack vessels in xylem.
- Phloem (वाही / पोषवाह): Conducts food materials from leaves to other parts.
- Elements: Sieve tube elements, Companion cells, Phloem parenchyma, Phloem fibers.
- Note: Gymnosperms have albuminous cells and sieve cells (no sieve tubes and companion cells).
- Xylem (दारू): Conducts water and minerals from roots to stem and leaves.
2. The Tissue System
Based on structure and location, plant tissue systems are of three types:
-
Epidermal Tissue System (बाह्यत्वचीय ऊतक तंत्र):
- Outer covering of the plant body.
- Includes epidermal cells, stomata, and epidermal appendages (trichomes and hairs).
- Stomata: Structures in epidermis of leaves regulating transpiration and gaseous exchange. Guard cells enclose the stomatal pore (bean-shaped in dicots, dumbbell-shaped in monocots).
-
Ground Tissue System (भरण ऊतक तंत्र):
- All tissues except epidermis and vascular bundles.
- Consists of parenchyma, collenchyma, and sclerenchyma.
- In leaves, the ground tissue is called Mesophyll (photosynthetic).
-
Vascular Tissue System (संवहन ऊतक तंत्र):
- Consists of Xylem and Phloem arranged in Vascular Bundles.
- Radial: Xylem and phloem arranged in alternate radii (e.g., Roots).
- Conjoint: Xylem and phloem situated along the same radius (e.g., Stems and Leaves).
- Open: Cambium present between xylem and phloem -> Secondary growth possible (Dicot stem).
- Closed: Cambium absent -> No secondary growth (Monocot stem).
3. Anatomy of Dicotyledonous and Monocotyledonous Plants
A. Dicot Root vs. Monocot Root
| Feature | Dicot Root | Monocot Root |
|---|---|---|
| Vascular Bundles | Usually 2 to 4 (Diarch to Tetrarch) | Polyarch (usually more than six) |
| Cambium | Developed later (Secondary growth occurs) | Absent (No secondary growth) |
| Pith | Small or inconspicuous | Large and well-developed |
B. Dicot Stem vs. Monocot Stem
| Feature | Dicot Stem | Monocot Stem |
|---|---|---|
| Vascular Bundles | Arranged in a ring | Scattered throughout the ground tissue |
| Bundle Sheath | Absent | Present (Sclerenchymatous) |
| Hypodermis | Collenchymatous | Sclerenchymatous |
| Secondary Growth | Present | Absent |
C. Dicot Leaf (Dorsiventral) vs. Monocot Leaf (Isobilateral)
| Feature | Dicot Leaf | Monocot Leaf |
|---|---|---|
| Stomata | More on lower epidermis (Hypostomatic) | Equal on both surfaces (Amphistomatic) |
| Mesophyll | Differentiated into Palisade and Spongy parenchyma | Undifferentiated (Uniform mesophyll) |
| Bulliform Cells | Absent | Present (help in curling of leaves to minimize water loss) |
4. Secondary Growth (द्वितीयक वृद्धि)
Increase in girth (diameter) of stems and roots, typical of dicots and gymnosperms. Driven by two lateral meristems:
-
Vascular Cambium:
- Forms a ring of meristematic tissue between xylem and phloem.
- Cuts off cells inwards towards the center to form Secondary Xylem (Wood).
- Cuts off cells outwards to form Secondary Phloem.
- Spring Wood / Early Wood: Lighter in color, vessels with wider cavities formed in spring.
- Autumn Wood / Late Wood: Darker in color, vessels with narrower cavities formed in winter.
- Annual Rings: Combination of spring and autumn wood of one year; used to estimate the age of a tree (Dendrochronology).
- Heartwood (Duramen): Central dark-colored dead wood, filled with resins/tannins; non-functional in water conduction, provides mechanical support.
- Sapwood (Alburnum): Peripheral light-colored living wood; actively conducts water and minerals.
-
Cork Cambium (Phellogen):
- Develops in the cortex region.
- Cuts off cells internally to form Secondary Cortex (Phelloderm).
- Cuts off cells externally to form Cork (Phellem).
- Periderm: Phellem (Cork) + Phellogen (Cork cambium) + Phelloderm (Secondary cortex).
- Bark: All tissues outside the vascular cambium (includes secondary phloem and periderm).
- Lenticels: Lens-shaped openings in the periderm that permit gaseous exchange between internal tissues and the atmosphere.